The story concerns the adventures of four youngsters from the kingdom of Fynn named Firion, Maria, Guy and Leon. Their parents are killed during an invasion by the army of The Emperor of Palamecia, who has summoned forth monsters from Hell in his quest to dominate the world. Fleeing the Emperor's monsters, the four are attacked and left for dead. Firion, Maria, and Guy are rescued by Princess Hilda of Fynn, who has established a rebel base in the nearby town of Altair. Eager to prove their value to the resistance movement, the three remaining youths undertake a variety of missions against Palamecia and join forces with a variety of allies not only to defeat the Emperor, but to locate Maria's missing brother Leon as well.
After Hilda asks Firion, Maria and Guy to go to Fynn to talk to her fiancee, Prince Scott, the party go to Fynn and they found out that Scott is in a secret place of the town. When the party finally finds him in a secret room in the town Pub, he is in the last moments of his life, but he is still able to tell the party that the defeat of Fynn was because of Count Borghen's betrayal. He also tells the party about his brother Gordon, telling them that he knows that Gordon has great strength. Scott then gives Firion his ring to bring back to Hilda and tells him that he will always love Hilda, using his last breath to say those words. The party returns to Altair and Firion gives the ring to Hilda; she recognizes it as Scott's ring and asks Firion to keep it, saying that is a ring from a brave man. After that she asks the party to go to Salamand to search for Mythril. Josef, a member of the rebellion was sent to search for it but Hilda hasn't heard anything from him since he left to go to Salamand. Hilda's right-hand man, Minwu also joins the party for this mission.
The party finds Josef, but he is unwilling to give any information, as the Palamecian empire have kidnapped Nelly, Josef's daughter, and Borghen has threatened to kill her if he helps the resistance. The party travels to the Semitt Falls where Nelly is held and frees her from the empire. After defeating one of the imperial Sergeants, they take the Mythril and make their way back to Altair.
After the weaponsmith Tobul crafts Mythril Equipment for the Resistance, the party is dispatched to Bafsk. The Palamecian Empire have enslaved the residents and made them build a powerful airship known as the Dreadnought. It was built under the watchful eye of an imperial known as the Dark Knight, however he has now been withdrawn since the empire lost the Mythril and has been replaced by the hapless Borghen. This gives the resistance the perfect opportunity to destroy the Dreadnought before it is finished. However, just before the party reach the massive airship from the Bafsk Sewers, the Dark Knight, who actually has not left Bafsk after all, appears and takes off with the Dreadnought before the party reaches it. The Dreadnought attacks the cities of Poft, Paloom, Gatrea and Altair, but miraculously the secret base at Altair is unharmed. A plan is formed to use the Sunfire from Kasuhan Keep but to enter, they need either the Goddess Bell or the voice of a Kashuan. Josef helps the party enter some snow caves with a snowcraft, and the party retrieves the bell which is located within. On the way out, Borghen attacks the party, and although he is defeated, he sends a boulder after the party. Josef holds back the boulder to allow the party to escape, but is crushed by the boulder, dying in the process.
With heavy hearts but renewed determination to avenge Josef, the party heads for the abandoned kingdom of Kashuan to retrieve the Sunfire. The party uses the Goddess Bell to infiltrate the Keep, where they meet with Gordon who helps them to locate Egil's Torch, the only vessel that they can use to carry the Sunfire. The party defeats a Red Soul for the torch and retrieve the Sunfire. As they leave, they witness Cid's Airship being abducted by the Dreadnought which then parks to replenish fuel supplies in the far south. The party heads there, frees Cid (and Hilda, who was on board Cid's Airship), and throws the Sunfire into the Dreadnought's engine as directed by Cid thus destroying it once and for all.
The party returns triumphantly to Altair, only to learn that the King is almost ready to die. With his last breaths, he forms a three-pronged attack on the Empire in an attempt to take back Fynn. In his plan, Minwu heads to Mysidia to retrieve Ultima, the ultimate magic tome; Gordon takes command of the rebel army to attack Fynn directly; and Firion's party heads to the island nation of Deist to enlist the aid of the Dragoons. With the help of Leila, the pirate captain, the party reaches Deist but finds that only a single Wyvern remains alive. However, the beast is dying, poisoned by the empire and it gives the party the last Wyvern Egg, which the party then drops in the healing spring at the bottom of Deist Cavern in order to incubate it and hasten its growth process.
The party returns to Altair empty-handed and to their surprise the Hilda they rescued on the Dreadnought is not really Hilda at all; it is a Lamia Queen that is impersonating the real Hilda. They soon learn that she is being held as a prize in the tournament at the Palamecian Coliseum. The party, with Gordon in tow, make haste to reach Hilda, defeating a Behemoth at the Coliseum to earn Hilda as a prize. However, The Emperor, who is overseeing the match personally, dispatches the party and has them thrown in the dungeons. They are saved by Paul the traveling thief, who unlocks their cell. Hilda and Gordon then escape on their own while the rest of the party draws the attention of the guards.
An attack is planned on Castle Fynn and the rebel army sets up just outside of town. Firion, Maria, Guy, and Leila lead the attack and the castle's incumbent, Gottos, is defeated, earning the rebels an important victory. However, Minwu and the Ultima Tome are nowhere to be found so Gordon instructs the party to look for Minwu and retrieve the Tome from the Mysidian Tower. After obtaining the Crystal Rod, the party heads for the Tower, only to be swallowed by Leviathan. Shipwrecked and without Leila, the party works its way from Leviathan's innards to the mouth, where, with the help of Ricard Highwind the Dragoon, they are able to retake the ship by defeating a Roundworm. They then head for the Tower, best the Fire Gigas, Ice Gigas and the Thunder Gigas which lie within and find Minwu at the Chamber of the Seal, desperately trying to break it to open the way to Ultima. In one last ditch effort, Minwu succeeds, but at a hefty price: he too succumbs to death to help the party in their battle against the Emperor.
The party takes Ultima and returns to Fynn but something is amiss. The towns of Altair, Gatrea, Paloom, and Poft have been destroyed by a mysterious force known as the Cyclone. It threatens to tear the world asunder if the party cannot figure out how to stop it. However, Gordon's idea of "sprouting wings" paves the way for the hatching of the last Wyvern, who comes to the castle to help the party reach the Cyclone.
Eventually, the party defeats the Emperor in the Cyclone. After the death of the Emperor, Leon, whom the party knew as the Emperor's Dark Knight and his right-hand man, decides to crown himself as the new Emperor. Firion and his party go to Palamecia to stop him, but when the party faces Leon, the Emperor comes back from Hell, more powerful than ever and the intention of reclaiming his Empire. Ricard Highwind stays in the castle of Palamecia to fight the Emperor, so that the party and Leon can escape from the castle on the Wyvern but the Emperor kills the Dragoon with ease. After the death of Ricard, the Dark Emperor raises Castle Pandaemonium, the fortress of the Lord of Hell, to start a new Empire. After returning to Deist to earn Excalibur, the treasured sword of the Dragoons, the party fights its way through the Jade Passage, entering Pandaemonium from underneath as all other forms of approach are impossible. Inside the castle, the party fights its way through several of the Emperor's most powerful minions, including the reincarnation of General Borghen himself, en route to the Emperor's throne at the top of the castle.
A fierce battle ensues as the Emperor attempts to destroy the last hope of the resistance. Despite his powerful spells and his ability to call down meteors, the Emperor eventually is at last defeated and dissipates into nothing, damned to the very Hell he directed against the party for so long. Firion and the party breathe a sigh of relief, and then return to Castle Fynn where Hilda, Gordon, Nelly, Leila, and Paul all wait to congratulate the party on the victory. In the aftermath of the battle, life begins anew for all these characters. However, Leon is skeptical of his own future since so much has gone on between the party members and him. Despite Maria's protests, Firion lets Leon go but reminds him that there is always a place for him there in Fynn, where he belongs.
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